Why ATP Yield from Glucose Varies
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One glucose molecule's ATP yield varies because the number of hydrogen ions the complexes pump differs between species, and the shuttle moving NADH into the mitochondrion differs between tissues.
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Liver cells shuttle glycolysis's NADH in using NAD+, while brain cells use a shuttle ending in FAD+, yielding fewer ATP.
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This range reflects real shuttle-system differences between cells, not an accounting error.