Male Gametophyte Development: Microsporogenesis and the Pollen Grain
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Inside an anther's microsporangium, mother cells divide by meiosis into microspores developing into pollen grains, each with a generative cell and a tube cell.
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Upon germination, a grain's tube cell forms the pollen tube while the generative cell divides into two sperm cells.
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The shed pollen grain lacks sperm; the generative cell forms two only as the tube grows toward the ovule.