Morgan's Drosophila Evidence for the Chromosomal Theory
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Thomas Hunt Morgan confirmed the chromosomal theory in 1910 by tracing a Drosophila eye-color trait's inheritance to the X chromosome specifically.
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Morgan's white-eye trait passed exactly as an X-linked factor would, never as an autosomal one.
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Morgan's result ties one named trait to one named chromosome — it is not itself proof every trait works this way.